What Is the Palisade Mesophyll and Its Function?

The palisade mesophyll is a specialized tissue found within plant leaves. It plays a significant role in the internal structure of a leaf, contributing to the plant’s ability to create its own food.

Where It’s Found and What It Looks Like

The palisade mesophyll is precisely positioned within a typical plant leaf, located directly beneath the upper epidermis and its protective waxy cuticle. This arrangement places it prominently in the upper part of the leaf, situated above the spongy mesophyll layer. It often forms a distinct layer or multiple layers.

Macroscopically, this layer contributes to the darker green appearance often seen on the upper surface of a leaf compared to its underside. Microscopically, the palisade mesophyll is characterized by its elongated, columnar, or cylindrical cells. These cells are tightly packed together, standing upright and typically arranged perpendicular to the leaf surface.

The Cells That Make It Up

The individual cells forming the palisade mesophyll exhibit specific characteristics. Their distinctive columnar or cylindrical shape allows them to be tightly packed, with minimal spaces between them. This dense arrangement helps to maximize the amount of tissue within a given leaf area.

A defining feature of these cells is their high concentration of chloroplasts, which are organelles containing the green pigment chlorophyll. Palisade cells contain the highest number of chloroplasts per cell compared to other plant tissues. These chloroplasts are capable of moving within the cell, adjusting their position to optimize light absorption.

Beyond chloroplasts, palisade cells also contain a large central vacuole, which occupies a significant portion of the cell volume. This vacuole helps maintain internal pressure and pushes chloroplasts towards the cell’s periphery, closer to the light source. A thin layer of cytoplasm surrounds the vacuole, containing other organelles like the nucleus and mitochondria. The cell is enclosed by a rigid cellulose cell wall and a cell membrane.

How It Powers Plant Life

The palisade mesophyll’s primary function is photosynthesis, the process by which plants convert light energy into chemical energy. This tissue is considered the main site of photosynthesis within the leaf. The structural features of palisade cells are specifically adapted to maximize the efficiency of this process.

The high density of chloroplasts within each palisade cell directly increases the capacity for light absorption. Their columnar shape and tight, upright packing minimize light scattering and allow for efficient light penetration deeper into the tissue. This arrangement ensures that a maximum surface area is exposed to incoming sunlight.

During photosynthesis, chlorophyll within the chloroplasts absorbs light energy, particularly from the blue and red regions of the light spectrum. Carbon dioxide, obtained from the atmosphere, enters the leaf through small pores called stomata and diffuses into the palisade cells. Water, absorbed by the roots, is transported to the leaves. These components are then used to produce glucose, a sugar that serves as the plant’s energy source, and oxygen, which is released as a byproduct into the atmosphere. The ability of the palisade mesophyll to efficiently carry out photosynthesis is fundamental for plant survival and contributes to the oxygen and food chains that sustain many life forms on Earth.