What Is an Incomplete Metamorphosis?

Insects undergo metamorphosis, a biological process of transformation that allows them to develop from an immature form to an adult. This remarkable change can follow one of two primary pathways: complete metamorphosis or incomplete metamorphosis. The latter, incomplete metamorphosis, involves a more gradual transition, setting it apart as a distinct developmental strategy in the insect world.

Understanding the Process

Incomplete metamorphosis, also known as hemimetabolism, describes a type of insect development where the changes from egg to adult are gradual rather than abrupt. This process is characterized by the absence of a pupal stage, which is a key feature of complete metamorphosis. Instead, the immature forms, called nymphs, generally resemble miniature versions of the adults, though they lack fully developed wings and reproductive organs.

Throughout their development, insects undergoing incomplete metamorphosis grow by shedding their outer skeleton, known as molting. Each stage between molts is referred to as an instar. The nymphs typically share similar habitats and diets with their adult counterparts, unlike the distinct ecological roles often seen in complete metamorphosis.

Developmental Stages

The life cycle of an insect with incomplete metamorphosis consists of three distinct stages: egg, nymph, and adult.

The life cycle begins with an egg, laid by the female insect. Nymphs hatch from these eggs. As they grow, nymphs periodically shed their exoskeleton through molting, with each stage between molts called an instar. Nymphs resemble miniature adults, though they lack fully developed wings and reproductive organs. They often share habitats and diets with adults.

After undergoing several molts, the nymph transitions into the adult stage. The adult insect is fully grown, possesses functional wings (if applicable to the species), and is reproductively mature. Once the adult stage is reached, no further molting occurs.

Distinguishing From Complete Metamorphosis

The primary distinction between incomplete and complete metamorphosis lies in the number of developmental stages and the appearance of the immature forms. Incomplete metamorphosis involves three stages: egg, nymph, and adult, while complete metamorphosis includes four stages: egg, larva, pupa, and adult.

A defining characteristic of incomplete metamorphosis is the absence of a pupal stage. Insects with complete metamorphosis undergo a pupal stage, a non-feeding, often inactive phase where significant transformation occurs. Their immature forms, larvae, are distinctly different from the adult. In contrast, nymphs resemble miniature adults. Larvae also frequently have different ecological roles and food sources than adults.

Insects That Undergo Incomplete Metamorphosis

Many familiar insects exhibit incomplete metamorphosis. Examples include grasshoppers and crickets, whose nymphs gradually develop wings through molts. Dragonflies and damselflies also follow this pattern; their aquatic nymphs, called naiads, live underwater and breathe with gills before emerging as winged adults.

Other insects that undergo incomplete metamorphosis include:
Cockroaches
True bugs (such as cicadas and aphids)
Praying mantises
Termites
Earwigs