What Are Cloacas and What Animals Have Them?

A cloaca represents a singular opening found in many animal species, serving as a versatile exit point for various bodily processes. Unlike most placental mammals, which possess distinct openings for waste elimination and reproduction, numerous other vertebrates utilize this common chamber.

What a Cloaca Is and Its Functions

A cloaca is a common internal chamber where the digestive, urinary, and reproductive tracts converge before opening to the outside. This single opening is responsible for the excretion of both solid waste, such as feces, and liquid waste, including urine.

Beyond waste expulsion, the cloaca also plays a central role in reproduction. It facilitates the passage of gametes, such as sperm and eggs, and is the opening through which eggs are laid or live young are born. In many birds, mating occurs through a brief contact known as a “cloacal kiss,” where sperm is transferred from the male to the female.

In some animals, the cloaca is internally divided into specialized compartments. For instance, in birds and reptiles, it typically includes the coprodeum for feces, the urodeum for urine and reproductive products, and the proctodeum, which is the final chamber before the external opening. Some species also use their cloaca for other purposes, such as scent marking through associated glands or even for respiration in certain aquatic turtles.

Animals That Possess a Cloaca

Many diverse groups within the animal kingdom possess a cloaca. This includes all amphibians, reptiles, and birds. The presence of a cloaca is considered a primitive feature in tetrapods, reflecting an ancient anatomical design.

All bird species have a cloaca, often referred to as a vent. During the breeding season, the cloacal regions of male and female birds often swell, enabling the transfer of sperm through direct contact.

Reptiles, including snakes, lizards, and turtles, universally feature a cloaca. Amphibians, such as frogs and salamanders, similarly utilize a cloaca.

Among fish, a true cloaca is present in cartilaginous fish like sharks and rays, as well as in lobe-finned fishes. However, most bony fishes do not have a cloaca, instead possessing separate openings.

Among mammals, only monotremes—the egg-laying mammals like the platypus and echidnas—possess a cloaca. The name “monotreme” itself derives from Greek words meaning “single opening,” directly referencing this anatomical characteristic. This distinguishes them from most other mammals, which have separate excretory and reproductive openings.