What Material Makes Up the Shell or Test of an Urchin?

Sea urchins are marine invertebrates known for their rigid outer covering, called a “test” or “shell.” This protective structure encases their soft internal organs, providing support and defense in diverse marine environments. Found in all oceans, these creatures have developed a durable casing. Understanding its composition reveals insights into natural biomineralization processes and how this material is intricately organized to create a robust skeletal system.

The Fundamental Material

The primary substance comprising a sea urchin’s test is calcium carbonate (CaCO3). This compound is present as calcite, a crystalline form. Calcite in sea urchin tests often contains a high content of magnesium, forming magnesium-rich calcite. This magnesium incorporation can contribute to the material’s properties, including its solubility and strength.

Sea urchins acquire the necessary components for shell formation directly from their surrounding seawater. They absorb calcium ions and convert dissolved carbon dioxide from the water into carbonate ions through a process involving specific enzymes, such as carbonic anhydrase. This biological conversion and deposition of calcium carbonate is a fundamental aspect of biomineralization, a process shared with other marine calcifiers like corals and mollusks. The resulting calcium carbonate provides structural integrity and protection against environmental challenges.

How Urchin Tests Are Built

The sea urchin test is not a single, continuous piece but a complex arrangement of numerous interlocking plates, known as ossicles. These plates are fused together, forming a rigid yet often spherical endoskeleton that grows with the animal. This modular design allows for growth without molting.

Each of these calcite plates possesses a unique internal architecture called stereom. Stereom is a spongy, lattice-like microstructure composed of microscopic calcitic trabeculae, which are small, rod-like structures. This intricate network makes the test porous and lightweight while maintaining considerable strength. The stereom essentially functions as a single crystal of calcite, despite its complex, fenestrated appearance.

An organic matrix, primarily composed of proteins and glycoproteins, is present within the mineral components of the test. This organic material plays a role in organizing the three-dimensional framework for mineralization and influences the properties of the biomineral. It helps bind the calcite crystals and contributes to the test’s overall resilience, allowing it to withstand significant pressure and impact by distributing force across its many plates.